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Taxidea
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« on: March 20, 2003, 11:43:31 AM »

 From the latest JHE (Volume 44, Issue 3 , March 2003,
 Pages 331-371

 Radiocarbon dating the appearance
 of modern humans and timing of
 cultural innovations in Europe: new
 results and new challenges

 Nicholas J. Conard,  and Michael Bolus

 Institut für Ur- und Frühgeschichte und Archäologie des
 Mittelalters, Universität Tübingen, Schloss
 Hohentübingen, 72070, Tübingen, Germany

 Received 1 February 2002;  revised 8 November 2002;  
 accepted 18 November 2002. ; Available
 online 19 March 2003.


    Abstract

    New radiocarbon dates from the sites of Bockstein-Törle,  
   Geißenklösterle, Hohle Fels,
    Hohlenstein-Stadel, Sirgenstein, and Vogelherd in the  
    Swabian Jura of southwestern Germany
    indicate that the Aurignacian of the region spans the
    period from ca. 40-30 ka BP. If the situation at Vogelherd,
   in which skeletal remains from modern humans underlie an  
   entire Aurignacian sequence, is viewed as representative for
  the region, the dates from the Swabian
    Jura support the hypothesis that populations of modern  
   humans entered the region by way of
    the "Danube Corridor." The lithic technology from the lower
   Aurignacian of Geißenklösterle III
    is fully developed, and classic Aurignacian forms are well  
   represented. During the course of
    the Aurignacian, numerous assemblages rich in art works,
   jewelry, and musical instruments are
    documented. By no later than 29 ka BP the Gravettian was
   well established in the region.
    These dates are consistent with the "Kulturpumpe"
   hypothesis that important cultural
    innovations of the Aurignacian and Gravettian in Swabia
   predate similar developments in other
    regions of Europe. The radiocarbon dates from
   Geißenklösterle corroborate observations from
    other non-archaeological data sets indicating large global
   fluctuations in the atmospheric
    concentrations of radiocarbon between 30 and 50 ka
   calendar years ago. These fluctuations
    lead to complications in building reliable chronologies during
   this period and cause the "Middle
    Paleolithic Dating Anomaly" and the "Coexistence Effect,"
   which tend to exaggerate the
    temporal overlap between Neanderthals and modern
   humans.

    Author Keywords: Neanderthals; Homo sapiens sapiens;
    Chronostratigraphy; Swabian
    Jura; Middle and Upper Paleolithic; Cultural innovations

 End abstract

 What does the last sentence of the abstarct imply? That
 the late Neanderthals are a fiction caused by faulty dating?
 Which, presumably, is the "Middle Paleolithic Dating
 Anomaly" or 'Coexistence effect'

 Comments or info anyone?

 Rick Wagler
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« Reply #1 on: March 22, 2003, 04:40:25 PM »

Quote
Quote from: Taxidea

From the latest JHE (Volume 44, Issue 3 , March 2003, Pages 331-371

Radiocarbon dating the appearanceof modern humans and timing of cultural innovations in Europe: new results and new challenges

Nicholas J. Conard,  and Michael Bolus
<snip>
Abstract:

New radiocarbon dates from the sites of Bockstein-Törle, Geißenklösterle, Hohle Fels, Hohlenstein-Stadel, Sirgenstein, and Vogelherd in the Swabian Jura of southwestern Germany indicate that the Aurignacian of the region spans the period from ca. 40-30 ka BP. If the situation at Vogelherd, in which skeletal remains from modern humans underlie an  entire Aurignacian sequence, is viewed as representative for the region, the dates from the Swabian Jura support the hypothesis that populations of modern humans entered the region by way of the "Danube Corridor." The lithic technology from the lower Aurignacian of Geißenklösterle III is fully developed, and classic Aurignacian forms are well represented. During the course of the Aurignacian, numerous assemblages rich in art works, jewelry, and musical instruments are documented. By no later than 29 ka BP the Gravettian was well established in the region. These dates are consistent with the "Kulturpumpe" hypothesis that important cultural innovations of the Aurignacian and Gravettian in Swabia predate similar developments in other regions of Europe. The radiocarbon dates from Geißenklösterle corroborate observations from other non-archaeological data sets indicating large global fluctuations in the atmospheric concentrations of radiocarbon between 30 and 50 ka calendar years ago. These fluctuations lead to complications in building reliable chronologies during this period and cause the "Middle Paleolithic Dating Anomaly" and the "Coexistence Effect," which tend to exaggerate the temporal overlap between Neanderthals and modern  humans.
<snip>

Quote

 What does the last sentence of the abstarct imply? That
 the late Neanderthals are a fiction caused by faulty dating?
 Which, presumably, is the "Middle Paleolithic Dating
 Anomaly" or 'Coexistence effect'

 Comments or info anyone?

 Rick Wagler

Since the word used by the authors is "exaggerate", I see no need to suggest that (or even to ask if), in their eyes, "the late Neanderthals are a fiction". This is, after all, just an Abstract which, by the way, seems to indicate that the actual article is clearly an update on a research issue that Conard & Bolus have been trying to deal with for a while. For example, see, (below), the abstract of a presentation that was made at the EUG XI MEETING (European Union of Geosciences) 8th - 12th April 2001, Strasbourg – France. Note also that, under whatever label, the chronological window under discussion is rather dirty, and this for all types of Quaternarists trying to study the events of the early and middle Interpleniglacial.
Quote

The Pattern and Timing of the Colonization of
Europe by Modern Humans
EUGXI.B3.2699
Nicholas J. Conard & Michael Bolus


Abundant fossil evidence indicates that modern humans evolved outside of Europe. Despite nearly a century and a half of research across many parts of the continent, there is no evidence for the arrival of anatomically modern humans in Europe before ca. 40 ka. At sites including Vogelherd where early European modern humans have been recovered, their skeletal remains are found in association with Aurignacian artifacts that are characterized by innovative lithic and organic artifacts and abundant examples of art and ornament that go far beyond the range of variation documented among the material culture of indigenous Neanderthals (Bolus & Conard, 2000; Churchill & Smith, 2000).

The distribution of dates for the most convincing evidence for the Aurignacian and early Upper Paleolithic suggests that Homo sapiens sapiens arrived in Europe around 40 ka. The best evidence for this early phase of occupation comes, broadly speaking, from the Circum-Mediterranean area and the upper reaches of the Danube Valley (Richter et al., 2000). The observation that many very early and also very rich Aurignacian sites are well documented in the Swabian Alb, suggests an early settlement of this area which led to the elaborate cultural developments documented at sites including Sirgenstein, Vogelherd, Hohlenstein-Stadel and Geißenklösterle. These sites, among other important finds, have yielded the earliest artworks and musical instruments known from Europe The Kulturpumpe model describes several potential mechanisms to explain these cultural innovations (Conard & Floss, 2000). Following this initial appearance of modern humans, Neanderthals and anatomically and culturally modern people coexisted in often spatial separate regions until ca. 25-30 ka when the last Neanderthals died out in the most peripheral parts of Europe.

This paper provides new dates for the Aurignacian of the Swabian Alb and archaeological analyses to test the hypotheses that the “Danube Corridor” and the Mediterranean coast provide the two key routes for the colonization of Europe. Ecological and chronological data testing the Kulturpumpe model will also be provided.


Jacques Cinq-Mars


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« Reply #2 on: March 24, 2003, 12:05:59 AM »




Note also that, under whatever label, the chronological window under discussion is rather dirty, and this for all types of Quaternarists trying to study the events of the early and middle Interpleniglacial.

Jacques Cinq-Mars



Some of this was discussed in the two free papers available from the previously-posted topic: "On the 'calibration' of radiocarbon dates"

CLICK HERE FOR THE URL

Also in:

Pettitt, P.B. (1999). Disappearing from the world: an archaeological perspective on Neanderthal extinction. Oxford Journal of Archaeology. 18 (3): 217-240.

the author cites "potential problems with 14C chronology" include the following (abstracted and abbreviated from pp. 224-225):

1) The asymptotic nature of C14 dates around 40ka BP. "...C14 ages appear to be asymptotic in nature, i.e. there is a 'C14 Event horizon' as Chappell et al. (1996) refer to it, around 40ka BP, at which residual contamination of samples is such that results tend to cluster around this period."

2) Correction of C14 dates beyond 10ka BP and age underestimation (= accuracy).  Here, Pettitt refers to the time of Neanderthal extinction as "the latter half of Oxygen Isotope Stage 3, a period within which atmospheric concentration of 14C seems to have been 20-40% higher than it is today and in which therefore all age determinations may well be underestimates (Duplessy 1996)....The underestimation of 14C between 25 and 50ka may be as much as 6ka, and according to some scholars, rises with age, reaching c. 10,000 years for samples at c. 50 ka BP (e.g. Geyh and Schlichter 1997; Goslar et al. 1997), although data are preliminary and there is, as yet, no general agreement; it may be that the offset decreases beyond c. 35kaBP (cf. Kitagawa and Van der Plicht 1998).  In all, we are in a period of some considerable flux with regards to such factors..."

3) Plateau in the 40-30ka BP time period and accuracy.  "...Perhaps the major problem affecting accuracy in the period concerned is the possibility of one or more plateaux between c. 40 and 30ka BP.  A potential plateau of major proportions is suggested by the fact that almost all speleothems with 'real' ages (i.e. dated by the other means than radiocarbon) of between 35 and 45ka date to c. 30,000 radiocarbon years BP (Goslar et al. 1997), an observation noted by Vogel (1983) where stalagmites dating to between 29 to 35ka cal BP (290Th) yielded 14C ages fairly consistent around 29,500 BP...."  Pettitt goes on to suggest other possible plateaux [at c. 32 and perhaps c. 35 ka], "...and for this reason alone one must accept that dates in the 30-40ka BP time period are provisional at best....Whatever the cause, it is clear that 14C dates at c. 31ka BP are some 5ka too young and that major radiocarbon plateaux probably exist in this broad period..."

Returning to the terms "Middle Paleolithic Dating Anomaly" and "Coexistence Effect", used by Conard & Bolus (2003) in their abstract, they seem to imply that some of the above "tend to exaggerate the temporal overlap between Neanderthals and modern humans".  Maybe they mean this could occur if the "C14 Event horizon" at 40 ka is used to suggest Neanderthal extinction began starting to occur at near that time, while the underestimation of dates of last occurrence, in some instances, might be as high as 5 ka (which would decrease the temporal overlap, effectively making the date of Neanderthal extinction earlier in these cases). This last paragraph, of course, is pure speculation on my part without access to the new paper by Conard & Bolus.

But clearly there are some problems with 14C dating in the period of 30-40 ka.

Dar  

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« Reply #3 on: March 24, 2003, 10:51:23 AM »

Quote from: Daryl Habel




Note also that, under whatever label, the chronological window under discussion is rather dirty, and this for all types of Quaternarists trying to study the events of the early and middle Interpleniglacial.

Jacques Cinq-Mars

Quote

Some of this was discussed in the two free papers available from the previously-posted topic: "On the 'calibration' of radiocarbon dates"

CLICK HERE FOR THE URL

Also in:...

<snip>

Returning to the terms "Middle Paleolithic Dating Anomaly" and "Coexistence Effect", used by Conard & Bolus (2003) in their abstract, they seem to imply that some of the above "tend to exaggerate the temporal overlap between Neanderthals and modern humans".  Maybe they mean this could occur if the "C14 Event horizon" at 40 ka is used to suggest Neanderthal extinction began starting to occur at near that time, while the underestimation of dates of last occurrence, in some instances, might be as high as 5 ka (which would decrease the temporal overlap, effectively making the date of Neanderthal extinction earlier in these cases). This last paragraph, of course, is pure speculation on my part without access to the new paper by Conard & Bolus.

But clearly there are some problems with 14C dating in the period of 30-40 ka.

Dar  


Thanks for the kind reminder. I suppose I should have mentioned some of this, especially since I am the one who posted the info about the Cologne Radiocarbon Calibration papers. Thanks also for the "clippings" from Pettitt's various statements on this issue. In this regard, I should add that Pettitt will have more to say on what Conard and Bolus call the "Middle Paleolithic Dating Anomaly" and "Coexistence Effect", in the Inaugural Issue of PALANTH which should be online in a few days!

Finally, I should also have mentioned that this particular debate is very much driven by the need on the part of palaeoanthropologists (and closely associated interdisciplinary colleagues), to arrive at a fairly high (level of chronological) resolution of the events they are interested in. This is not necessarily a concern shared by all the Quaternarists I was referring to.

Jacques
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« Reply #4 on: March 24, 2003, 05:20:47 PM »


From the latest JHE (Volume 44, Issue 3 , March 2003,
Pages 331-371

Radiocarbon dating the appearance
of modern humans and timing of
cultural innovations in Europe: new
results and new challenges

Nicholas J. Conard,  and Michael Bolus

Institut für Ur- und Frühgeschichte und Archäologie des
Mittelalters, Universität Tübingen, Schloss
Hohentübingen, 72070, Tübingen, Germany

Received 1 February 2002;  revised 8 November 2002;  
accepted 18 November 2002. ; Available
online 19 March 2003.


   Abstract

   New radiocarbon dates from the sites of Bockstein-Törle,  
  Geißenklösterle, Hohle Fels,
   Hohlenstein-Stadel, Sirgenstein, and Vogelherd in the  
   Swabian Jura of southwestern Germany
   indicate that the Aurignacian of the region spans the
   period from ca. 40-30 ka BP. If the situation at Vogelherd,
  in which skeletal remains from modern humans underlie an  
  entire Aurignacian sequence, is viewed as representative for
 the region, the dates from the Swabian
   Jura support the hypothesis that populations of modern  
  humans entered the region by way of
   the "Danube Corridor." The lithic technology from the lower
  Aurignacian of Geißenklösterle III
   is fully developed, and classic Aurignacian forms are well  
  represented. During the course of
   the Aurignacian, numerous assemblages rich in art works,
  jewelry, and musical instruments are
   documented. By no later than 29 ka BP the Gravettian was
  well established in the region.
   These dates are consistent with the "Kulturpumpe"
  hypothesis that important cultural
   innovations of the Aurignacian and Gravettian in Swabia
  predate similar developments in other
   regions of Europe. The radiocarbon dates from
  Geißenklösterle corroborate observations from
   other non-archaeological data sets indicating large global
  fluctuations in the atmospheric
   concentrations of radiocarbon between 30 and 50 ka
  calendar years ago. These fluctuations
   lead to complications in building reliable chronologies during
  this period and cause the "Middle
   Paleolithic Dating Anomaly" and the "Coexistence Effect,"
  which tend to exaggerate the
   temporal overlap between Neanderthals and modern
  humans.

   Author Keywords: Neanderthals; Homo sapiens sapiens;
   Chronostratigraphy; Swabian
   Jura; Middle and Upper Paleolithic; Cultural innovations

End abstract

What does the last sentence of the abstarct imply? That
the late Neanderthals are a fiction caused by faulty dating?
Which, presumably, is the "Middle Paleolithic Dating
Anomaly" or 'Coexistence effect'

Comments or info anyone?

Rick Wagler


Hi Rick,

2 comments.

First

It appears obvious to us that the Danube corridor is an potentially easy migration route into the heart of Europe.  I suspect that it would not necessarily have that appearance to a native African, who would see nearly any major river drainage as a potential tsetse habitat and even might see it as a deadly avenue to follow for that reason.   In most of Africa, nearly all river corridors are certain to be completely blocked in places by tsetse infestation.  I suspect that ease of movement between areas outside of Africa was much greater than between areas inside Africa, resulting in greater isolation of individual populations in Africa, therefore resulting in accelerated genetic drift in those populations.  That would "exaggerate" the appearance of age in the characters of those distinct populations mostly likely to be subject to that degree and type of isolation.  

Second

Are there not quite late Neanderthal examples in Croatia, late enough even with "Middle Paleolithic Dating Anomaly" mentioned to put recognized populations of Neanderthals essentially in the same river corridor in very similar time frames to the non-Neanderthal populations in the Swabia?

Dutch
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« Reply #5 on: March 25, 2003, 03:41:24 AM »



2 comments.

First

It appears obvious to us that the Danube corridor is an potentially easy migration route into the heart of Europe.  I suspect that it would not necessarily have that appearance to a native African, who would see nearly any major river drainage as a potential tsetse habitat and even might see it as a deadly avenue to follow for that reason.   In most of Africa, nearly all river corridors are certain to be completely blocked in places by tsetse infestation.  I suspect that ease of movement between areas outside of Africa was much greater than between areas inside Africa, resulting in greater isolation of individual populations in Africa, therefore resulting in accelerated genetic drift in those populations.  That would "exaggerate" the appearance of age in the characters of those distinct populations mostly likely to be subject to that degree and type of isolation.



Hi Dutch,

The Danube is potentially an easy migration route into the heart of Europe, but it is not necessarily the only route to consider.  Conard and Bolus (2003, abstract) are advocating that, "innovations of the Aurignacian and Gravettian in Swabia predate similar developments in other regions of Europe," citing dates from Geissenklosterle.  Recent dating at Geissenklosterle (Richter et al. 2000) placed the earliest Aurignacian (level III at Geissenklosterle) in the Hengelo Interstadial (c. 43.0-40.5 kyr BP) by paleoecological analysis. 14C-AMS results yielded a mean of 38.4 +/- 0.85 kyr BP, whereas a mean of 40.2 +/- 1.5 kyr BP was obtained by TL on burnt silex, confirming the correlation of the level III Early Aurignacian with the Hengelo Interstadial.

The Swabian caves might be earliest manifestations in the "Danube Corridor" of the Aurignacian "innovations" as Conard & Bolus say, but they lie near the source of the river and far upstream of Bacho Kiro and Temnata, proposed by Bolus & Conard (2001:31) as "Early Balkan Aurignacian" entry point for the "replacement" migration.

But it could be questioned whether the Aurignacian "innovations" found in the Swabian caves actually predate similar developments at Kostenki?  Certainly Kostenki, on the East European Plain, lies in one of the "other regions of Europe" (Conard & Bolus 2003, abstract).   Hoffecker et al. (2002) write: "...A volcanic ash layer dated to 38,000-35,000 years BP represents a widespread stratigraphic marker horizon. Several Upper paleolithic occupation levels underlie this horizon at Kostenki 12 and 14, and they appear to date to roughly 45,000-40,000 years BP..."

In the press release from University of Colorado at Boulder (21 March 2002), Hoffecker is quoted: "The Kostenki sites, which date beyond 40,000 years ago, may have hosted Neanderthals as well as modern humans...It looks like there were two separate industries at work here.  One culture was advanced in terms of bone and ivory tool-making and decorative figurine art, while the other produced little more than crude stone tools".  

Regardless of Hoffecker's opinion of the Streletskaya industry (the one with the "crude stone tools"),  both it and the "advanced" (usually described as the Aurignacoid technocomplex) lie in the "lower humic beds" at Kostenki, and have been correlated (by A.A. Sinitsyn) with the Hengelo Interstadial.

So if we have an industry with the same basic "innovations" (bone and ivory tool making and decorative figurine art) at Geissenklosterle and Kostenki, and both are dated (paleoecologically) to the Hengelo Interstadial, what would be the logical "corridor" connecting these two localities?

I would suggest we not forget the Moravian Gate (SW of Krakow, Poland) which was certainly in use for crossing the Carpathians from Eastern Europe by the later Gravettian populations.  I'm not advocating anything specific here (dating is inadequate, and it's a long, long way from G to K), but there are possible migration routes into the upper Danube which do not require traveling the lower river.


Quote


Second

Are there not quite late Neanderthal examples in Croatia, late enough even with "Middle Paleolithic Dating Anomaly" mentioned to put recognized populations of Neanderthals essentially in the same river corridor in very similar time frames to the non-Neanderthal populations in the Swabia?

Dutch


Bolus & Conard  (2001:31) say of this:  "The exact stratigraphic positions and cultural attributions, as well as the dates of the hominid remains from Southwestern Central Europe belonging to this period, have been questioned (Mellars et al. 1999; Zilhao and d'Errico, 1999a, b).  Among others, this is true for the Neanderthals from the Early Upper Paleolithic level G1 of Vindija cave in Croatia (Fig. 1:11) which yielded AMS radiocarbon ages of 33 +/- 0.4 kyr BP (Karavanic and Smith, 1998, Fig. 2) and 28-29 kyr BP (Smith et al. 1999) respectively..."

Although they are not critical "in the specifics", there does seem, in this wording, to be an inclination towards questioning the dating of the Croatian Neanderthals.  Taken along with their (Conard & Bolus 2003, abstract) expressed idea of a possible tendency for dating effects to "exaggerate" AMH/N temporal overlap, I might guess that any possible reduction of this overlap would better fit their "Kulturpumpe" hypothesis, but I've not looked into this far enough to be sure.

Peace back at ya,
Dar
 

References:

Bolus, M. & Conard, N.J. (2001). The late Middle Paleolithic and earliest Upper Paleolithic in Central Europe and their relevance for the Out of Africa hypothesis. Quaternary International 75: 29-40.  

Hoffecker, J.F., Anikovich, M.V., Sinitsyn, A.A., Holliday, V.T. & Forman, S.L. (2002). Initial Upper Paleolithic in Eastern Europe: new research at Kostenki. Abstracts from the Paleoanthropology Society Meetings. Denver Colorado, March 19-20, 2002.

Richter, D., Waiblinger, J., Rink, W.J. & Wagner, G.A. (2000). Thermoluminescence, electron spin resonance and 14C-dating of the Late Middle and Early Upper Palaeolithic site of Geissenklostere Cave in Southern Germany. Journal of Archaeological Science 27: 71-89.


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« Reply #6 on: March 25, 2003, 10:54:11 AM »

Hi Dar,

I was wondering if the authors weren't pointing toward redating the Croatian Neanderthals, but in so doing they also push the Swabian cites back as well.  Or is it vise versa...  That would still have the dates for the two types more or less overlapping;  so I don't necessarily see that any relative interval was increased.

You are very correct about multiple possible corredors, which have to be considered to have run both ways.  It is certainly likely that at any particular point in time the main flow was in a single direction, but part of the story concerning Neanderthals and the succeeding graciles has to be who was traveling in which direction and when, going back much farther than any gracile presence in Swabia.

The Danube corredor is likely to have been more often available for those movements than those farther north during Ice Age climatic cycling in Europe.   I was impressed by topographical maps that it seems such a natural highway cutting through the heart of Europe that I cannot see that it was not used "going downhill" long before any documented gracile presence.  That would have put European Neanderthals at least irregularly into the Middle East during the very periods where we find a highly variable human population there.  I am more convinced than ever that Europe exported Neanderthal genetics into that area and did so consistantly.  At some point that downhill movement changed to uphill contaminating the Neanderthal core gene pool effectively ending the Neanderthal consistancy necessary for maintenance of type.  Or so it seems to me.

I was actually keying on two concepts in the abstract the first is dealt with above, the second was use of river corredors for human movement generally.  Or rather that humans in Africa could not have used them in the same way as they were used elsewhere, since they almost all have locally intense tsetse infestations in Africa, barring the kind of easy human mobility that they provide elsewhere.  It was "exaggerated temporal effects" that caught my attention.  The inhibition of the same kinds of river corredor traffic in Africa that we expect in the rest of the world is an important variable in the consideration of African genetic isolation.  Increased population isolation in Africa automatically supports the maintenance of increased local lineages, thus exaggerating appearance of their antiquity.  Instead of producing underrepresentation of actual time as was proposed for Europe, the effect in Africa would be overestimation of actual time elapsed, as more effectively isolated populations were better able to maintain distinctive diversity over the same periods than more mobile populations who would integrate.   This type of genetic isolation directly questions the accuracy of time tables attached to mtDNA trees native to Africa and considered to be close to basal in the African Eve scenario, since it is unaccounted for and obviously intense.  It is almost a certainty to my way of thinking that many African populations and their genetics could not possibly have participated in any Upper Paleolithic exodus, and may well end up being as external to current non-African populations even the Neanderthals.  Time lines and basal branching points badly need to be recalculated.

Sorry I don't have any scientific references to point you to.  There don't seem to be many in regard to specific barriers to human movement in Africa, although I did stumble on one in regard to the timing of the initial presence of domestic dogs in South Africa which mentioned east African migrations south as following tsetse free corredors along the Great Rift valley.  Not scientifically published, but in connection with the "africanis" or native dog type in Africa.

http://www.everythingdoggy.com/rareafricanus.htm

It mentions relatively slow Bantu migrations south through tsetse free corridors, seeming to assume that is common knowlege.  (that migration also includes the introduction of most other domesticates and Iron Age technology as well as the dog)  There must be something else.

One other peripheral bit may be that the original Islamic conversion movement stalled on the borders of tsetse country in Africa, which would be expected if important mobility considerations were imbedded in native cultures.  Islamic conversion involves a very complete discard of previous folklores, which may well have resulted in the loss of important survival behaviours.  

Personally I expect that the movement folklores are very subtle in native African cultures.  Since they involve a factor completely alien to European experience, and a zoonotic factor not well understood by anyone until recently, they may have gone unnoticed, but it strikes me for example that a native African herdsman would be very reluctant to follow common American practice and regularly use gallery and bottom land for pasturage for his cattle and probably has dozens of "old wives tales" about why and how to avoid natural dangers in that practise in his native areas.  

I would be grateful for any pointers or references about native social control of movement in Africa.  From what I have seen so far I would expect them to show very careful movement of livestock and location of mother-child reproductive units and cropping areas as well as an enforced general delay in male reproductive participation until a maturing adolescent can show particular levels of skill in avoiding dangers in longer range mobility.  IOW a quite intensive localization of the human reproductive and economic bases in Africa beyond that inherant anywhere else in the world quite simply due to presence of the tsetse fly which would make these considerations very, very ancient, potentially as as ancient as the Homo presence in Africa and perhaps even older.

BTW I have been unable to find references about Pan tolerance to tsetse vectored tyrpanosomes either, which may be an important consideration to divergence at LCA.  Based on their native range, I would expect Pan to show the tolerance that other African non-human primates exhibit. which humans do not share, but that remains a question mark to me, as it does for Gorilla.   That tolerance is documented for vervets and baboons and IIRC African colubus types.  The lack of that tolerance is definitely documented and absolute in humans.   So where does that change?

It is good to reason with you again.  As you can see I need help with some points, either to develop them or to put them to sleep for good.  Back to the drawing board is not a very big step at this point and I would rather take it sooner than later should it be necessary. <grin>

Peace

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